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The families of flowering plants.                                                                                                                                                                

Tribelaceae (Engl.) Airy Shaw

                        

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~Escalloniaceae, Grossulariaceae sensu lato

Habit and leaf form. Prostrate shrubs (the robust, flexuous stems bearing short, ascending leafy branches); not resinous. Leaves alternate; thickish, glaucescent above; sessile (semiamplexicaul); non-sheathing; simple. Lamina entire; ‘obscurely nerved’. Leaves exstipulate. Lamina margins apically minutely tri- dentate.

Leaf anatomy. Lamina without secretory cavities.

Stem anatomy. Secretory cavities absent. Secondary thickening developing from a conventional cambial ring (?).

Reproductive type, pollination. Unisexual flowersabsent. Plants hermaphrodite.

Inflorescence, floral, fruit and seed morphology. Flowerssolitary (at the apices of branchlets); regular; 5 merous; cyclic; tetracyclic. Free hypanthium absent. Hypogynous disk absent.

Perianthwith distinct calyx and corolla; 10; 2 whorled; isomerous. Calyx 5 (small); 1 whorled; shortly gamosepalous (below); regular; persistent; imbricate. Corolla 5 (relatively large); 1 whorled; polypetalous; contorted; regular; fleshy to not fleshy (‘thickish’). Petals slightly clawed; entire (elliptical).

Androecium 5. Androecial members free of the perianth; all equal; free of one another; 1 whorled. Androecium exclusively of fertile stamens. Stamens5; isomerous with the perianth; oppositisepalous; alternating with the corolla members; filantherous (the filaments subulate). Anthers not becoming inverted during development; dehiscing via longitudinal slits; small, ovoid, extrorse.

Gynoecium 3 carpelled. Carpels reduced in number relative to the perianth. The pistil3 celled. Gynoecium syncarpous; synstylovarious to eu-syncarpous; superior. Ovary 3 locular. Gynoecium stylate. Styles 1 (short, simple); apical. Stigmas 1; 3 lobed; capitate. Placentation axile. Ovules 30–50 per locule (‘many’); anatropous.

Fruit non-fleshy (the epicarp leathery, the endocarp crustaceous); dehiscent; a capsule. Capsules loculicidal and valvular (borne on a short, erect pedicel, the valves finally separating from the axis, to which the seeds long remain attached after the capsule has dehisced). Fruit many seeded. Seedscopiously endospermic. Endosperm oily. Testa smooth; black.

Geography, cytology. Neotropical and Antarctic. Pampas and Andean. Temperate. Temperate South America.

Taxonomy.Subclass Dicotyledonae; Tenuinucelli (?). Dahlgren’s Superorder Corniflorae (?); Cornales (?). Cronquist’s Subclass Rosidae; Rosales. APG 3 core angiosperms; core eudicot; unplaced at Superordinal level (?); Order Escalloniales (as a synonym of Escalloniaceae?).

Species 1. Genera 1; only genus, Tribeles.

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