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The families of flowering plants.                                                                                                                                                                

Heteropyxidaceae Engler & Gilg

                        

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~Myrtaceae

Habit and leaf form. Small trees. Leaves alternate; spiral; petiolate; non-sheathing; gland-dotted (pellucid punctate); simple. Lamina entire; pinnately veined (without an intramarginal vein); cross-venulate. Leaves stipulate. Lamina margins entire. Domatia occurring in the family; manifested as pockets.

Leaf anatomy. Lamina with secretory cavities.

Stem anatomy. Internal phloem present (?). Secondary thickening developing from a conventional cambial ring. Xylem with fibre tracheids; without libriform fibres.

Reproductive type, pollination. Plants hermaphrodite.

Inflorescence, floral, fruit and seed morphology. Flowers aggregated in ‘inflorescences’; in panicles. Inflorescences terminal; terminal thyrses. Flowers small; regular; 5 merous; cyclic; tetracyclic. Free hypanthium present (the corolla ‘inserted on the calyx tube’).

Perianth with distinct calyx and corolla; 10; 2 whorled; isomerous. Calyx 5; 1 whorled; polysepalous, or gamosepalous; five blunt-lobed; regular; persistent; scarcely imbricate, or open in bud. Corolla 5; 1 whorled; polypetalous (the small petals inserted on the hypanthium/calyx tube). Petals shortly clawed (and gland dotted).

Androecium 5. Androecial members free of the perianth (inserted on the tube); all equal; free of one another; 1 whorled. Androecium exclusively of fertile stamens. Stamens 5; isomerous with the perianth; alternisepalous; opposite the corolla members; inflexed in bud. Anthers dorsifixed; dehiscing via longitudinal slits; introrse; appendaged (with a well developed connective gland). The anther appendages apical. Pollen grains aperturate; 3 aperturate; (parasyn-) colporate.

Gynoecium2–3 carpelled. Carpels reduced in number relative to the perianth. The pistil 2–3 celled. Gynoecium syncarpous; eu-syncarpous; superior. Ovary 2–3 locular; sessile. Gynoecium stylate. Styles 1; from a depression at the top of the ovary; apical; short, persistent. Stigmas 1; capitate. Placentation axile (the ovules over most of the peltate placenta). Ovules30–50 per locule (‘many’, no ovulodes); hemianatropous, or campylotropous (?).

Fruit non-fleshy; dehiscent; a capsule. Capsules loculicidal (small, 2–3 locular). Seedsnon-endospermic. Cotyledons 2; flat. Embryo straight.

Geography, cytology. Paleotropical. Tropical. South Africa.

Taxonomy.Subclass Dicotyledonae; Crassinucelli. Dahlgren’s Superorder Myrtiflorae; Myrtales. Cronquist’s Subclass Rosidae; Myrtales. APG 3 core angiosperms; core eudicot; Superorder Rosanae; malvid; Order Myrtales (as a synonym of Myrtaceae).

Species 3. Genera 1; only genus, Heteropyxis.

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